(This variant rule is inspired by predator behaviour in Neanderthal).
Creatures in Bios: Megafauna can expand to an apex predator slot if they meet the following requirements:
1. They are able to predate the carnivore in their home biome.
2. They are able to predate at least 2 carnivores in adjacent locations, provided they can move to them.
Adjacent apex predators compete if they have any prey in common - even if they do not depend on this prey in order to be supported. Competition is resolved using the carnivore competition rules.
Note: Apex predators depend primarily on scavenging kills from carnivores, rather than directly hunting.
The apex predator slot may be expanded to or entered if a species has nowhere else available to occupy.