I Am One With The Force
The Force Is With Me
The official ruling is to go according to what the thing you're playing on says. It's more fun that way, too -- adds variety.
Agreed. I didnt even notice the Villains mat was different until I played using Burrow for the first time.
Now selecting to use the board or the mat is just one of the setup variables I take into consideration when playing.
Alternatively, if you want to use the mat but using the Hero direction, just move Villain deck to the escape pile and move left to right instead of right to left.